Who invented karyotype?
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Lev Delaunay in 1922 seems to have been the first person to define the karyotype as the phenotypic appearance of the somatic chromosomes, in contrast to their genic contents.
Cyril Darlington pioneered plant cytogenetics in 1920–30 and made important advances in our understanding of mechanisms of chiasma formation and the behavior of sex chromosomes in meiosis